OK. There's no hardware reason why the extruder motors won't turn. So I still think it's a software problem. Marlin is working since the other motors work, so I think Marlin is intentionally stopping you from extruding. Look at the log/terminal window -- it should be telling you why.Maybe the thermistor is shorted so it's seeing a MAXTEMP rather than a low temperature? I don't know if this applies to Marlin, but Repetier checks that the temperature is increasing when the heater is on, and shuts down if not. Once an error is detected, you need to send an M999 to re-enable things.
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